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FOSSILS TELL OUR STORY
ARCHAEOLOGY TELLS OUR STORY
DNA TELLS OUR STORY
Denisova Cave, Siberia
Paabo Svante & a Neanderthal friend
Homo heidelbergensis in relation to Neanderthals and Sapiens
Australopithecus
Australopithecus is a group of extinct primates closely related to, if not actually ancestors of, modern human beings and known from a series of fossils found at numerous sites in eastern, north-central, and southern Africa. The various species of Australopithecus lived 4.4 million to 1.4 million years ago (mya), during the Pliocene and Pleistocene epochs (which lasted from 5.3 million to 11,700 years ago). The genus name, meaning “southern ape,” refers to the first fossils found, which were discovered in South Africa. Perhaps the most famous specimen of Australopithecus is “Lucy,” a remarkably preserved fossilized skeleton from Ethiopia that has been dated to 3.2 mya. As characterized by the fossil evidence, members of Australopithecus bore a combination of humanlike and apelike traits. They were similar to modern humans in that they were bipedal (that is, they walked on two legs), but, like apes, they had small brains. Their canine teeth were smaller than those found in apes, and their cheek teeth were larger than those of modern humans.
Homo erectus
Early African Homo erectus fossils (sometimes called Homo ergaster) are the oldest known early humans to have possessed modern human-like body proportions with relatively elongated legs and shorter arms compared to the size of the torso. These features are considered adaptations to a life lived on the ground, indicating the loss of earlier tree-climbing adaptations, with the ability to walk and possibly run long distances. Compared with earlier fossil humans, note the expanded braincase relative to the size of the face. The most complete fossil individual of this species is known as the ‘Turkana Boy’ – a well-preserved skeleton (though minus almost all the hand and foot bones), dated around 1.6 million years old. Microscopic study of the teeth indicates that he grew up at a growth rate similar to that of a great ape. There is fossil evidence that this species cared for old and weak individuals. The appearance of Homo erectus in the fossil record is often associated with the earliest handaxes, the first major innovation in stone tool technology. Early fossil discoveries from Java (beginning in the 1890s) and China (‘Peking Man’, beginning in the 1920s) comprise the classic examples of this species. Generally considered to have been the first species to have expanded beyond Africa, Homo erectus is considered a highly variable species, spread over two continents (it's not certain whether it reached Europe), and possibly the longest lived early human species - about nine times as long as our own species, Homo sapiens, has been around (i.e. nearly 1.8 to 2.0 million years)!
Turkana Boy (AMNH)
Homo Neanderthalensis
Neanderthals are an extinct group of archaic hominins who lived in Eurasia until about 40,000 years ago. The type specimen, Neanderthal 1, was found in 1856 in the Neander Valley in present-day Germany. It is not clear when the line of Neanderthals split from that of modern humans; studies have produced various times ranging from 315,000 to more than 800,000 years ago. The date of divergence of Neanderthals from their ancestor H. heidelbergensis is also unclear. The oldest potential Neanderthal bones date to 430,000 years ago, but the classification remains uncertain. Neanderthals are known from numerous fossils, especially from after 130,000 years ago. The reasons for Neanderthal extinction are disputed. Neanderthals lived in a high-stress environment with high trauma rates; about 80% of Neanderthal individuals died before the age of 40. The total population of Neanderthals remained low (less than 75 thousand), and interbreeding with modern humans tended toward a loss of Neanderthal genes over time. They lacked effective long-distance networks. Despite this, there is evidence of regional cultures and regular communication between communities, possibly moving between caves seasonally. For much of the early 20th century, European researchers depicted Neanderthals as primitive, unintelligent and brutish. Although knowledge and perception of them has markedly changed since then in the scientific community, the image of the unevolved caveman archetype remains prevalent in popular culture. Compared with modern humans, Neanderthals had a more robust build and proportionally shorter limbs. Researchers often explain these features as adaptations to conserve heat in a cold climate; however, they may also have been adaptations for sprinting in the warmer, forested landscape that Neanderthals often inhabited. They had cold-specific adaptations, such as specialized body-fat storage. Average Neanderthal men stood around 165 cm (5 ft 5 in) and women 153 cm (5 ft 0 in) tall, similar to pre-industrial modern Europeans. The braincases of Neanderthal men and women averaged about 1,600 cm3 (98 cu in) and 1,300 cm3 (79 cu in), respectively, which is considerably larger than the modern human average. Although they were probably apex predators, they still competed with cave lions, cave hyenas, and other large predators. A number of examples of symbolic thought and Paleolithic art have been attributed to Neanderthals. Some claims of religious beliefs have been made. Neanderthals were likely capable of speech, possibly articulate, although the complexity of their language is not known. The 2010 Neanderthal genome project's draft report presented evidence for interbreeding between Neanderthals and modern humans. Neanderthals also appear to have interbred with Denisovans, a different group of archaic humans, in Siberia. Around 1–4% of genomes of Eurasians, Indigenous Australians, Melanesians, Native Americans and North Africans is of Neanderthal ancestry, while most inhabitants of sub-Saharan Africa have around 0.3% of Neanderthal genes, save possible traces from early sapiens-to-Neanderthal gene flow and/or more recent back-migration of Eurasians to Africa. In all, about 20% of distinctly Neanderthal gene variants survive in modern humans. Neanderthal introgression appears to have affected the modern human immune system and is also implicated in several other biological functions and structures, including the ear and nasal cavities.
Homo Sapiens
Humans (Homo sapiens) or modern humans are the most common and widespread species of primate, and the last surviving species of the genus Homo. They are great apes characterized by their hairlessness, bipedalism, and high intelligence. Humans have large brains, enabling more advanced cognitive skills that enable them to thrive and adapt in varied environments, develop highly complex tools, and form complex social structures and civilizations. Humans are highly social, with individual humans tending to belong to a multi-layered network of distinct social groups – from families and peer groups to corporations and political states. As such, social interactions between humans have established a wide variety of values, social norms, languages, and traditions (collectively termed institutions), each of which bolsters human society. Humans are also highly curious: the desire to understand and influence phenomena has motivated humanity's development of science, technology, philosophy, mythology, religion, and other frameworks of knowledge; humans also study themselves through such domains as anthropology, social science, history, psychology, and medicine. As of 2025, there are estimated to be more than 8 billion humans alive. For most of their history, humans were nomadic hunter-gatherers. Humans began exhibiting behavioral modernity about 160,000–60,000 years ago. The Neolithic Revolution, which began in Southwest Asia around 13,000 years ago (and separately in a few other places), saw the emergence of agriculture and permanent human settlement; in turn, this led to the development of civilization and kickstarted a period of continuous (and ongoing) population growth and rapid technological change. Since then, a number of civilizations have risen and fallen, while a number of sociocultural and technological developments have resulted in significant changes to the human lifestyle.
Although the term "humans" technically equates with all members of the genus Homo, in common usage it generally refers to Homo sapiens, the only extant member. All other members of the genus Homo, which are now extinct, are known as archaic humans, and the term "modern human" is used to distinguish Homo sapiens from archaic humans. Anatomically modern humans emerged around 300,000 years ago in Africa, evolving from Homo heidelbergensis or a similar species. Migrating out of Africa, they gradually replaced and interbred with local populations of archaic humans. Multiple hypotheses for the extinction of archaic human species such as Neanderthals include competition, violence, interbreeding with Homo sapiens, or inability to adapt to climate change. Genes and the environment influence human biological variation in visible characteristics, physiology, disease susceptibility, mental abilities, body size, and life span. Though humans vary in many traits (such as genetic predispositions and physical features), humans are among the least genetically diverse primates. Any two humans are at least 99% genetically similar. Humans are sexually dimorphic: generally, males have greater body strength, and females have a higher body fat percentage. At puberty, humans develop secondary sex characteristics. Females are capable of pregnancy, usually between puberty, at around 12 years old, and menopause, around the age of 50. Humans are omnivorous, capable of consuming a wide variety of plant and animal material and have used fire and other forms of heat to prepare and cook food since the time of Homo erectus. Humans have had a dramatic effect on the environment. They are apex predators, being rarely preyed upon by other species. Human population growth, industrialization, land development, overconsumption, and combustion of fossil fuels have led to environmental destruction and pollution that significantly contribute to the ongoing mass extinction of other forms of life. Within the last century, humans have explored challenging environments such as Antarctica, the deep sea, and outer space, though human habitation in these environments is typically limited in duration and restricted to scientific, military, or industrial expeditions. Humans have visited the Moon and sent human-made spacecraft to other celestial bodies. Humans can survive for up to eight weeks without food and several days without water. Humans are generally diurnal, sleeping on average seven to nine hours per day. Childbirth is dangerous, with a high risk of complications and death. Often, both the mother and the father provide care for their children, who are helpless at birth.